diff --git a/django/core/management/__init__.py b/django/core/management/__init__.py
index 6edb0ba..16b27c8 100644
--- a/django/core/management/__init__.py
+++ b/django/core/management/__init__.py
@@ -140,9 +140,12 @@ def call_command(name, *args, **options):
     # when the script runs from the command line, but since call_command can
     # be called programatically, we need to simulate the loading and handling
     # of defaults (see #10080 for details).
-    defaults = dict([(o.dest, o.default)
-                     for o in klass.option_list
-                     if o.default is not NO_DEFAULT])
+    defaults = {}
+    for opt in klass.option_list:
+        if opt.default is not NO_DEFAULT:
+            defaults[opt.dest] = opt.default
+        else:
+            defaults[opt.dest] = None
     defaults.update(options)
 
     return klass.execute(*args, **defaults)
diff --git a/tests/modeltests/user_commands/management/commands/dance.py b/tests/modeltests/user_commands/management/commands/dance.py
index 34eb277..4ad5579 100644
--- a/tests/modeltests/user_commands/management/commands/dance.py
+++ b/tests/modeltests/user_commands/management/commands/dance.py
@@ -9,8 +9,10 @@ class Command(BaseCommand):
     requires_model_validation = True
 
     option_list =[
-        make_option("-s", "--style", default="Rock'n'Roll")
+        make_option("-s", "--style", default="Rock'n'Roll"),
+        make_option("-x", "--example")
     ]
 
     def handle(self, *args, **options):
+        example = options["example"]
         self.stdout.write("I don't feel like dancing %s." % options["style"])
